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i <br />Z <br />mortality, food habits, and habitat suitability were collected for a sample of 174 <br />nests. Family size averaged 3.0 individuals per group with 630/0 of the breeding <br />pairs accompanied by at least one helper. Clutch size averaged 3.8 and average <br />brood size was 3.0 (Toland unpubl. data). Median incubation date was April 7'h <br />and median hatching date was April 24"'. Approximately 72% of the nestlings <br />successfully fledged. Mean annual productivity was 1.9 young fledged per pair <br />per year. Median fledging date was May 10"' and 70% of all nesting attempts <br />were successful in fledging at least one young (Toland unpubl. data). <br />Habitat characteristics were assessed in one-half acre patches around each nest <br />tree and categorized as optimal, suboptimal, ancillary, or lawn (Toland 1991). <br />Nesting success (percentage of nest attempts that fledged at least one young) was <br />positively correlated to nest site habitat quality; success rates for optimal, <br />suboptimal, ancillary, and lawn were 91, 67, 48, and 25%, respectively. Habitat <br />features comprising each scrub -jay territory were assessed and characterized as <br />optimal contiguous, optimal fragmented, and suboptimal fragmented. Nesting <br />success was positively correlated with territory quality; success rates for optimal <br />contiguous, optimal fragmented, and suboptimal fragmented were 79, 77, and <br />59%, respectively (Toland 1991). Mean annual productivity of scrub -jays in the <br />study area was 2.2 young fledged per pair in contiguous, optimal scrub; 1.8 young <br />fledged per pair in fragmented, moderately developed scrub; 1.2 young per pair <br />fledged in highly fragmented, suboptimal scrub; and about 0.5 young per pair in <br />residential lawns. tong -term studies at Archbold Biological Station have previously <br />documented that mean annual productivity of scrub jays is 2,0 young per pair per <br />year in stable populations where natality counterbalances mortality (Fitzpatrick et <br />a/. 1991). <br />The subject demographic studies found that scrub jays dispersed at earlier ages and <br />farther distances than is reported for contiguous natural scrub landscapes (Toland <br />unpubl. data), A total of 64 dispersals were documented, including 40 females and <br />24 males (Toland unpubl. data). The mean dispersal distance by females was 2.4 <br />miles (3.8 km) lrange=0.2 mi. (0.3 kin) to 5 mi. (8.0 kin)] and the mean dispersal <br />distance by males was 0.9 miles (1.4 km) (range -0.6 mi. (0.075 kin) to 2.9 mi. <br />(4.7 km)]. Four females were known to disperse 4 miles (6.5 km) between the <br />most northern clusters of the north subpopulation and the Sebastian <br />Highlands/Pelican Island Elementary School southernmost clusters (Toland unpubl. <br />data). The longest dispersal was 5 miles from a female that emigrated into the <br />Wabasso Scrub Conservation Area frorn a cluster located on the Sebastian <br />Airport/Golf Course (Toland unpubl. data). Most of the documented dispersals in <br />the City of Sebastian were 2 miles or less (80% of females and 92% of males) <br />(Toland unpubl. data). <br />Breininger (1999) docurnented similar patterns on the urbanized and extensively <br />fragmented South Brevard County barrier island, where females dispersed an <br />average of 4.5 miles (7.3 km) and reales dispersed an average of 0.6 miles (1.0 <br />37 <br />